What structure of the sarcomere shortens during muscle contraction?

The Z lines and H- zone shortens during the muscle contraction.

Explanation:

Sarcomere is the area between the two Z lines. The Z- lines are present in the centre of I - band and H - zone is present in the A - band. The actin filaments which are thin filaments present in I- band and contracts, slide over the myosin filament during contraction.

The I - band, H- zones are became shortens as they are thin than the A- band. It is anisotropic band having thick myosin filaments. They are not flexible and remain in its constant shape during the muscle contraction.

After the muscle contraction, the A- band and the H- zone comes to their original shape. In other words, the sarcomere shortens and comes back during muscle contraction, and relaxation.

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the answer to your question is:

c. complementary base-pairing