In which population was natural selection not the reason for the difference in fitness?

Although the operation of natural selection requires that genotypes differ in fitness, for some geneticists it seems easier to understand natural selection than fitness. Partly this reflects the fact that the word “fitness” has been used to mean subtly different things. Here I distinguish among these meanings (e.g., individual versus absolute versus relative fitness) and explain how evolutionary geneticists use fitness to predict changes in the genetic composition