Malthus formed his theory by studying factors that control the population growth of humans. how might factors operating on organisms in nature differ from those of malthus's theory?

This is like a humans vs organisms in nature. Consider what makes humans different from any other living organism.

Factors that control human population growth:

food
shelter
goods
water
disease
health
relationships

Factors that control nature population growth:

- food
- shelter
- precipitation
- relationships
- food web
- weather ( natural disasters)

So as you can see humans pretty much has more control over our population growth while nature operates by cycle and a set of systems that man can't create, but can manipulate.

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It is used to during sexual reproduction between humans