The fuzzy millipede, Polyxenus fasciculatus, is found in Texas and is preyed upon by most species of ants. To protect itself against the ants, it ejects fibers from a tuft located at the tail end of its body. The tufts have hooks at the tips and barbs along their length that lock and interlink with the ant’s setae, small hairs that cover the body of the ant. When an ant attacks, the millipede flexes its back end toward the ant and wipes the tufts against it. As the ant attempts to remove the tufts, it entangles itself more, becoming immobilized. Which two systems most directly interact in the fuzzy millipede’s defense against ants?