Fossils of a whale that beached on an african shore more than a million years ago and was subsequently butchered by hominids have been recovered by paleontologists.

a. that beached on an african shore more than a million years ago and was subsequently butchered by hominids have

b. that beached on an african shore more than a million years ago and then was subsequently butchered by hominids has

c. that beached on an african shore more than a million years ago, which was subsequently butchered by hominids, has

d. having been beached on an african shore more than a million years ago and subsequently butchered by hominids, have

e. having beached on an african shore more than a million years ago and then subsequently were butchered by hominids have