An article's subject may influence

when it is published
whether is effectively received
who reads it
what the writer intends to do for the reader
all of the above

C) who reads it. 
The audience is determined by the subject, and who reads it is the audience. Hope this helps!

The best and most correct answer among the choices provided by your question is all of the above which is the fifth choice.
An article's subject may influence:

when it is published
whether is effectively received
who reads it
what the writer intends to do for the reader

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